The revolution that still isn't: The origins of behavioral complexity in Homo sapiens.

The behavioral origins of Homo sapiens can be traced back to the first material culture produced by our species in Africa, the Middle Stone Age (MSA). Beyond this broad consensus, the origins, patterns, and causes of behavioral complexity in modern humans remain debated. Here, we consider whether recent findings continue to support popular scenarios of: (1) a modern human 'package,' (2) a gradual and 'pan-African' emergence of behavioral complexity, and (3) a direct connection to changes in the human brain. Our geographically structured review shows that decades of scientific research have continuously failed to find a discrete threshold for a complete 'modernity package' and that the concept is theoretically obsolete. Instead of a continent-wide, gradual accumulation of complex material culture, the record exhibits a predominantly asynchronous presence and duration of many innovations across different regions of Africa. The emerging pattern of behavioral complexity from the MSA conforms to an intricate mosaic characterized by spatially discrete, temporally variable, and historically contingent trajectories. This archaeological record bears no direct relation to a simplistic shift in the human brain but rather reflects similar cognitive capacities that are variably manifested. The interaction of multiple causal factors constitutes the most parsimonious explanation driving the variable expression of complex behaviors, with demographic processes such as population structure, size, and connectivity playing a key role. While much emphasis has been given to innovation and variability in the MSA record, long periods of stasis and a lack of cumulative developments argue further against a strictly gradualistic nature in the record. Instead, we are confronted with humanity's deep, variegated roots in Africa, and a dynamic metapopulation that took many millennia to reach the critical mass capable of producing the ratchet effect commonly used to define contemporary human culture. Finally, we note a weakening link between 'modern' human biology and behavior from around 300 ka ago.

Materiality, Agency and Evolution of Lithic Technology: an Integrated Perspective for Palaeolithic Archaeology.

Considerations of materiality and object-oriented approaches have greatly influenced the development of archaeological theory in recent years. Yet, Palaeolithic archaeology has been slow in incorporating this emerging body of scholarship and exploring its bearing on the human deep past. This paper probes into the potential of materiality theory to clarify the material dynamics of the Plio-Pleistocene and seeks to re-articulate the debate on the evolution of our species with materiality discourses in archaeology and the humanities more broadly. We argue that the signature temporalities and geospatial scales of observation provided by the Palaeolithic record offer unique opportunities to examine the active role of material things, objects, artefacts and technologies in the emergence, stabilisation and transformation of hominin lifeworlds and the accretion of long-term trajectories of material culture change. We map three axes of human-thing relations-ecological, technical and evolutionary-and deploy a range of case studies from the literature to show that a critical re-assessment of material agency not only discloses novel insights and questions, but can also refine what we already know about the human deep past. Our exploration underscores the benefits of de-centring human behaviour and intentionality and demonstrates that materiality lends itself as a productive nexus of exchange and mutual inspiration for diverging schools and research interests in Palaeolithic archaeology. An integrated object-oriented perspective calls attention to the human condition as a product of millennial-scale human-thing co-adaptation, in the course of which hominins, artefacts and technologies continuously influenced and co-created each other.

Different environmental variables predict body and brain size evolution in Homo.

Increasing body and brain size constitutes a key macro-evolutionary pattern in the hominin lineage, yet the mechanisms behind these changes remain debated. Hypothesized drivers include environmental, demographic, social, dietary, and technological factors. Here we test the influence of environmental factors on the evolution of body and brain size in the genus Homo over the last one million years using a large fossil dataset combined with global paleoclimatic reconstructions and formalized hypotheses tested in a quantitative statistical framework. We identify temperature as a major predictor of body size variation within Homo, in accordance with Bergmann's rule. In contrast, net primary productivity of environments and long-term variability in precipitation correlate with brain size but explain low amounts of the observed variation. These associations are likely due to an indirect environmental influence on cognitive abilities and extinction probabilities. Most environmental factors that we test do not correspond with body and brain size evolution, pointing towards complex scenarios which underlie the evolution of key biological characteristics in later Homo.

Patterns of extreme temperature-related catastrophic events in Europe including the Russian Federation: a cross-sectional analysis of the Emergency Events Database.

OBJECTIVE: To investigate reported extreme temperature-related catastrophic events and associated mortality on the European continent including the Russian Federation. DESIGN: Cross-sectional respecting Strengthening the Reporting of Observational Studies in Epidemiology (STROBE) criteria. SETTINGS: Data source: Emergency Events Database (EM-DAT). PARTICIPANTS: Search criteria: location-European continent including Russian Federation, time-years 1988 until 2019 (close of database 12 July 2019), catastrophic events-extreme temperatures. PRIMARY OUTCOME MEASURES: Numbers of heat waves, cold waves, severe winter conditions and associated number of deaths, overall, and per country and year, respecting STROBE criteria. RESULTS: The most frequent type of the 243 events recorded in EM-DAT were cold waves (54.7%). However, cold waves and severe winter conditions only accounted for 6460 deaths (4.5%), while heat waves were associated with 137 533 deaths (95.5%). The five most severe heat waves in 2003, 2006, 2010, 2013 and 2015 were associated with a total of 135 089 deaths. The most severe heat waves were geographically distributed over the Russian Federation (2010), as well as France, Italy, Spain and Germany, each in 2003. CONCLUSION: Although cold waves are more frequently reported in EM-DAT, heat waves are the major cause for temperature-related deaths. In order to better protect the public, it is important to address resiliency and vulnerability of populations at risk and age groups.

Regional patterns of diachronic technological change in the Howiesons Poort of southern Africa.

The Howiesons Poort (HP) of southern Africa plays an important role in models on the early behavioral evolution of Homo sapiens. The HP is often portrayed as a coherent MSA industry characterized by early complex material culture. Recent work has emphasized parallel technological change through time across southern Africa potentially driven by ecological adaptations or demographic change. Here we examine patterns of diachronic variation within the HP and evaluate potential causal factors behind these changes. We test previous temporal assessments of the technocomplex at the local and regional level based on high-resolution quantitative data on HP lithic assemblages from Sibudu (KwaZulu-Natal) and comparisons with other southern African sites. At Sibudu, consistent unidirectional change in lithic technology characterizes the HP sequence. The results show a gradual reduction in typical HP markers such as the proportion of blades, backed pieces, and HP cores, as well as declining size of blades and backed artifacts. Quantitative comparisons with seven HP sites in South Africa suggest that lithic technology varies between regions over time instead of following similar changes. Concerning hypotheses of causal drivers, directional changes in lithic technology at Sibudu covary with shifting hunting patterns towards larger-sized bovids and a gradual opening of the vegetation. In contrast, variation in lithic technology shows little association with site use, mobility patterns or demographic expansions. Unlike at Sibudu, diachronic changes at other HP sites such as Diepkloof, Klasies River and Klipdrift appear to be associated with aspects of mobility, technological organization and site use. The regional diachronic patterns in the HP partly follow paleoclimatic zones, which could imply different ecological adaptations and distinct connection networks over time. Divergent and at times decoupled changes in lithic traits across sites precludes monocausal explanations for the entire HP, supporting more complex models for the observed technological trajectories.

Long-term patterns of body mass and stature evolution within the hominin lineage.

Body size is a central determinant of a species' biology and adaptive strategy, but the number of reliable estimates of hominin body mass and stature have been insufficient to determine long-term patterns and subtle interactions in these size components within our lineage. Here, we analyse 254 body mass and 204 stature estimates from a total of 311 hominin specimens dating from 4.4 Ma to the Holocene using multi-level chronological and taxonomic analytical categories. The results demonstrate complex temporal patterns of body size variation with phases of relative stasis intermitted by periods of rapid increases. The observed trajectories could result from punctuated increases at speciation events, but also differential proliferation of large-bodied taxa or the extinction of small-bodied populations. Combined taxonomic and temporal analyses show that in relation to australopithecines, early Homo is characterized by significantly larger average body mass and stature but retains considerable diversity, including small body sizes. Within later Homo, stature and body mass evolution follow different trajectories: average modern stature is maintained from ca 1.6 Ma, while consistently higher body masses are not established until the Middle Pleistocene at ca 0.5-0.4 Ma, likely caused by directional selection related to colonizing higher latitudes. Selection against small-bodied individuals (less than 40 kg; less than 140 cm) after 1.4 Ma is associated with a decrease in relative size variability in later Homo species compared with earlier Homo and australopithecines. The isolated small-bodied individuals of Homo naledi (ca 0.3 Ma) and Homo floresiensis (ca 100-60 ka) constitute important exceptions to these general patterns, adding further layers of complexity to the evolution of body size within the genus Homo. At the end of the Late Pleistocene and Holocene, body size in Homo sapiens declines on average, but also extends to lower limits not seen in comparable frequency since early Homo.

Implications of Nubian-Like Core Reduction Systems in Southern Africa for the Identification of Early Modern Human Dispersals.

Lithic technologies have been used to trace dispersals of early human populations within and beyond Africa. Convergence in lithic systems has the potential to confound such interpretations, implying connections between unrelated groups. Due to their reductive nature, stone artefacts are unusually prone to this chance appearance of similar forms in unrelated populations. Here we present data from the South African Middle Stone Age sites Uitpanskraal 7 and Mertenhof suggesting that Nubian core reduction systems associated with Late Pleistocene populations in North Africa and potentially with early human migrations out of Africa in MIS 5 also occur in southern Africa during early MIS 3 and with no clear connection to the North African occurrence. The timing and spatial distribution of their appearance in southern and northern Africa implies technological convergence, rather than diffusion or dispersal. While lithic technologies can be a critical guide to human population flux, their utility in tracing early human dispersals at large spatial and temporal scales with stone artefact types remains questionable.

Examining the Causes and Consequences of Short-Term Behavioral Change during the Middle Stone Age at Sibudu, South Africa.

Sibudu in KwaZulu-Natal (South Africa) with its rich and high-resolution archaeological sequence provides an ideal case study to examine the causes and consequences of short-term variation in the behavior of modern humans during the Middle Stone Age (MSA). We present the results from a technological analysis of 11 stratified lithic assemblages which overlie the Howiesons Poort deposits and all date to ~58 ka. Based on technological and typological attributes, we conducted inter-assemblage comparisons to characterize the nature and tempo of cultural change in successive occupations. This work identified considerable short-term variation with clear temporal trends throughout the sequence, demonstrating that knappers at Sibudu varied their technology over short time spans. The lithic assemblages can be grouped into three cohesive units which differ from each other in the procurement of raw materials, the frequency in the methods of core reduction, the kind of blanks produced, and in the nature of tools the inhabitants of Sibudu made and used. These groups of assemblages represent different strategies of lithic technology, which build upon each other in a gradual, cumulative manner. We also identify a clear pattern of development toward what we have previously defined as the Sibudan cultural taxonomic unit. Contextualizing these results on larger geographical scales shows that the later phase of the MSA during MIS 3 in KwaZulu-Natal and southern Africa is one of dynamic cultural change rather than of stasis or stagnation as has at times been claimed. In combination with environmental, subsistence and contextual information, our high-resolution data on lithic technology suggest that short-term behavioral variability at Sibudu can be best explained by changes in technological organization and socio-economic dynamics instead of environmental forcing.

Spatial and temporal variation of body size among early Homo.

The estimation of body size among the earliest members of the genus Homo (2.4-1.5Myr [millions of years ago]) is central to interpretations of their biology. It is widely accepted that Homo ergaster possessed increased body size compared with Homo habilis and Homo rudolfensis, and that this may have been a factor involved with the dispersal of Homo out of Africa. The study of taxonomic differences in body size, however, is problematic. Postcranial remains are rarely associated with craniodental fossils, and taxonomic attributions frequently rest upon the size of skeletal elements. Previous body size estimates have been based upon well-preserved specimens with a more reliable species assessment. Since these samples are small (n < 5) and disparate in space and time, little is known about geographical and chronological variation in body size within early Homo. We investigate temporal and spatial variation in body size among fossils of early Homo using a 'taxon-free' approach, considering evidence for size variation from isolated and fragmentary postcranial remains (n = 39). To render the size of disparate fossil elements comparable, we derived new regression equations for common parameters of body size from a globally representative sample of hunter-gatherers and applied them to available postcranial measurements from the fossils. The results demonstrate chronological and spatial variation but no simple temporal or geographical trends for the evolution of body size among early Homo. Pronounced body size increases within Africa take place only after hominin populations were established at Dmanisi, suggesting that migrations into Eurasia were not contingent on larger body sizes. The primary evidence for these marked changes among early Homo is based upon material from Koobi Fora after 1.7Myr, indicating regional size variation. The significant body size differences between specimens from Koobi Fora and Olduvai support the cranial evidence for at least two co-existing morphotypes in the Early Pleistocene of eastern Africa.

Characterizing the Late Pleistocene MSA Lithic Technology of Sibudu, KwaZulu-Natal, South Africa.

Studies of the African Middle Stone Age (MSA) have become central for defining the cultural adaptations that accompanied the evolution of modern humans. While much of recent research in South Africa has focused on the Still Bay and Howiesons Poort (HP), periods following these technocomplexes were often neglected. Here we examine lithic assemblages from Sibudu that post-date the HP to further the understanding of MSA cultural variability during the Late Pleistocene. Sibudu preserves an exceptionally thick, rich, and high-resolution archaeological sequence that dates to ∼ 58 ka, which has recently been proposed as type assemblage for the "Sibudan". This study presents a detailed analysis of the six uppermost lithic assemblages from these deposits (BM-BSP) that we excavated from 2011-2013. We define the key elements of the lithic technology and compare our findings to other assemblages post-dating the HP. The six lithic assemblages provide a distinct and robust cultural signal, closely resembling each other in various technological, techno-functional, techno-economic, and typological characteristics. These results refute assertions that modern humans living after the HP possessed an unstructured and unsophisticated MSA lithic technology. While we observed several parallels with other contemporaneous MSA sites, particularly in the eastern part of southern Africa, the lithic assemblages at Sibudu demonstrate a distinct and so far unique combination of techno-typological traits. Our findings support the use of the Sibudan to help structuring this part of the southern African MSA and emphasize the need for further research to identify the spatial and temporal extent of this proposed cultural unit.

Coastal adaptations and the Middle Stone Age lithic assemblages from Hoedjiespunt 1 in the Western Cape, South Africa.

New excavations at the Middle Stone Age (MSA) open-air site of Hoedjiespunt 1 (HDP1) on the west coast of South Africa advance our understanding of the evolution of coastal adaptations in Homo sapiens. The archaeological site of HDP1 dates to the last interglacial and consists of three phases of occupation, each containing abundant lithic artifacts, shellfish, terrestrial fauna, ostrich eggshell and pieces of ground ocher. The site provides an excellent case study to analyze human behavioral adaptations linked to early exploitation of marine resources. Here we reconstruct human activities through a detailed study of the lithic assemblages, combining analyses of the reduction sequences, artifact attributes and quartz fracturing. These methods provide insights into raw material procurement, lithic reduction sequences, site use and mobility patterns, and foster comparison with other MSA coastal sites. The main characteristics of the lithic assemblages remain constant throughout the use of the site. Quartz dominates silcrete and other raw materials by almost four to one. Knappers at HDP1 produced different forms of flakes using multiple core reduction methods. Denticulates represent the most frequent tool type. The assemblages document complete, bipolar and hard hammer reduction sequences for the locally available quartz, but highly truncated reduction sequences with many isolated end products for silcrete, a material with a minimum transport distance of 10-30km. This observation suggests that well provisioned individuals executed planned movements to the shoreline to exploit shellfish. Our excavations at HDP1 furthermore demonstrate the simultaneous occurrence of flexible raw material use, anticipated long-distance transport, systematic gathering of shellfish and use of ground ocher. The HDP1 lithic assemblages document a robust pattern of land-use that we interpret as a stable adaptation of modern humans to coastal landscapes as early as MIS 5e.